Mikoto is more reluctant than the other two Princesses to cross-dress because he does not want his girlfriend Megumi to find out about it. As such he is nonetheless a reliable character—he often stubbornly whines and complains about things the Princesses have to do but always ends up doing them anyway whether he's forced to or not. He is very self-conscious about being a Princess and is mortified whenever someone else outside of school finds out about it. He is the most animated of the three Princesses, always showing a lot of emotion and expression when yelling at Toru and Yuujirou. Mikoto is also tone deaf, having never sung before becoming a princess.
He also cares a lot about the people close to him, though especially for his girlfriend. He has shown to change character and act different when around Megumi, which greatly amuses Toru and Yuujirou. Despite all the whining and complaining, however, he eventually becomes accustomed to his duty as a Princess such that he once had a brief fantasy of him and Megumi both dressed in Gothic Lolita Princess.
He is usually in the Princess outfit, appearing as a nurse, Gothic Lolita Princess, etc. While he is cross dressing as a Princess, he wears a long wavy pink wig that matches his hair color. His wig is the only one that is wavy. On other days, he wears his school uniform, a coat-like uniform that is black and has a silver lined opening. His hair is short and pink.
Mikoto is dedicated to his girlfriend Megumi, to the extent that even his personality and character changes when around Megumi. He holds her in high respect, and does not want to embarrass himself in front of her. He enjoys her company very much, and takes her words seriously. Megumi is the only one that can calm Mikoto fully when he is embarrassed at her finding out that he is a cross-dressing Princess.
Mikoto sees Tooru as a good friend, but is annoyed by the fact that even though Tooru was given a chance to opt out of the Princess System, Tooru agreed readily when he heard about the benefits. He is even more frustrated that Tooru was even given a chance to opt out in the first place. When Tooru willingly gets into the role of becoming a Princess, Mikoto blames him and claims that he has gone over to the dark side, joining Yuujirou.
Mikoto has a love-hate relationship with Yuujirou. Though he acknowledges the fact that Yuujirou is a good friend, he is annoyed by how Yuujirou enjoys being a Princess. When he complains about having to cross-dress, he gets frustrated when Yuujirou taunts him. On impulse, he once claimed that Yuujiruo hated him, but seemed shocked by his own claim after he said it. He finds Yuujirou irritating, but an invaluable friend.